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ÐÎÑÑÈÉÑÊÀß ÀÊÀÄÅÌÈß ÍÀÓÊ ÐÓÑÑÊÎÅ ÝÍÒÎÌÎËÎÃÈ×ÅÑÊÎÅ ÎÁÙÅÑÒÂÎ
×òåíèÿ ïàìÿòè Íèêîëàÿ Àëåêñàíäðîâè÷à Õîëîäêîâñêîãî Âûï. 53
Ã.Ñ. Ìåäâåäåâ
Ýâîëþöèÿ è ñèñòåìà æóêîâ-÷åðíîòåëîê òðèáû Blaptini (Coleoptera, Tenebrionidae)
Ñàíêò-Ïåòåðáóðã 2001
Ìåäâåäåâ Ã.Ñ. Ýâîëþöèÿ è ñèñòåìà æóêîâ-÷åðíîòåëîê òðèáû Blaptini (Coleoptera, Tenebrionidae). ×òåíèÿ ïàìÿòè Í.À. Õîëîäêîâñêîãî. Âûï. 53, ÑÏá., 2001. 332 ñ. Medvedev G.S. Evolution and system of darkling beetles of the tribe Blaptini (Coleoptera, Tenebrionidae). Meetings in memory of N.A. Cholodkovsky. Iss. 53. StPetersburg, 2001. 332 pp.
Ðåäàêòîð Ñ.Þ.Ñèí¸â Îòâåòñòâåííûé çà èçäàíèå Â.À. Êðèâîõàòñêèé
Ïî ïîñòàíîâëåíèþ Ïðåçèäèóìà Ðîññèéñêîé àêàäåìèè íàóê åæåãîäíî â ìàðòå-àïðåëå ïðîâîäÿòñÿ ×òåíèÿ ïàìÿòè âûäàþùåãîñÿ ðóññêîãî çîîëîãà, ïî÷åòíîãî ÷ëåíà Ðóññêîãî ýíòîìîëîãè÷åñêîãî îáùåñòâà, ïðîôåññîðà Íèêîëàÿ Àëåêñàíäðîâè÷à Õ î ë î ä ê î â ñ ê î ã î (1858-1921). Íàñòîÿùèé âûïóñê ñîäåðæèò ðàñøèðåííîå èçëîæåíèå äîêëàäà Ã.Ñ. Ìåäâåäåâà, âûñòóïèâøåãî íà ïðîâîäèâøèõñÿ 31 ìàðòà 2000 ã. 53-õ ×òåíèÿõ ñ ñîîáùåíèåì î ìîðôîãåíåòè÷åñêèõ ïðåîáðàçîâàíèÿõ ðÿäà ïðèçíàêîâ ïîëîâîé ñèñòåìû è ãåíèòàëèé æóêîâ-÷åðíîòåëîê òðèáû Blaptini (Coleoptera, Tenebrionidae). Ïóáëèêàöèÿ âûïóñêà ïîääåðæàíà ãðàíòàìè ÐÔÔÈ 98-04-49763 è ÔÖÍÒÏ “Áèîðàçíîîáðàçèå”.
ISSN 1606-8858
© Ðóññêîå ýíòîìîëîãè÷åñêîå îáùåñòâî, 2001 © Ã.Ñ. Ìåäâåäåâ, 2001
Ã.Ñ. Ìåäâåäåâ Ñèñòåìà è ýâîëþöèÿ æóêîâ–÷åðíîòåëîê òðèáû Blaptini (Coleoptera, Tenebrionidae) Ñîäåðæàíèå Ââåäåíèå................................................................................................................4 1.Ìîðôîëîãèÿ......................................................................................................5 1.1. Îáùàÿ õàðàêòåðèñòèêà òðèáû .......................................................5 1.2. Ìóñêóëàòóðà ãîëîâû ...................................................................7 1.3. Ñòðîåíèå âåðøèíû áðþøêà ñàìöà ..............................................8 1.4. Ñòðîåíèå âåðøèíû áðþøêà ñàìêè ............................................10 2. Ïîëîæåíèå òðèáû Blaptini â ñèñòåìå ñåì. Tenebrionidae ..............................11 3. Ñèñòåìàòè÷åñêèé îáçîð ïîäòðèá ................................................................15 3.1. Ïîäòðèáà Blaptina Latreille, 1817 ................................................15 3.2. Ïîäòðèáà Remipedellina Semenow, 1906 ...................................21 3.3. Ïîäòðèáà Prosodina Skopin, 1960 ..............................................23 3.4. Ïîäòðèáà Gnaptorina G.Medvedev, subtrib. n. ...........................29 3.5. Ïîäòðèáà Gnaptorinina G.Medvedev, subtrib. n. ........................31 4. Îïðåäåëèòåëüíàÿ òàáëèöà ïîäòðèá è ðîäîâ ..............................................33 5. Æèçíåííûå ñòðàòåãèè ÷åðíîòåëîê òðèáû Blaptini ....................................39 6. Ìîðôî-ýâîëþöèîííûå òåíäåíöèè .............................................................42 6.1. Óñèêè ............................................................................................42 6.2. Ýäåàãóñ..........................................................................................45 6.2.1. Ôàëëîáàçà .....................................................................46 6.2.2. Ïàðàìåðû ......................................................................51 6.3. Ãàñòðàëüíàÿ ñïèêóëà ....................................................................61 6.4. ßéöåêëàä ......................................................................................67 6.5. Âåíòðàëüíàÿ ñïèêóëà ...................................................................68 6.6. Ïîëîâûå ïóòè ñàìêè ....................................................................70 7. Èçîëèðóþùèå ìåõàíèçìû ó ÷åðíîòåëîê ðîäà Prosodes ...........................79 7.1. Õèìè÷åñêàÿ èçîëÿöèÿ ............................................